Move with WASD / arrow keys (or the on-screen joystick on mobile). Aim with the mouse (or drag the right side of the screen on mobile) and hold Left Mouse to fire — auto-fire is on by default. Defeat hostile cells to drop plasma motes that magnet toward you and fill your mutation meter; when it fills, choose a branch, then stack upgrades. Press Space (or the on-screen button) for your branch ability once you reach Tier 2. Clear every chamber and elite to advance through six biozones to the Apex Core. Esc pauses, M mutes, F is fullscreen. Progress auto-saves in your browser.
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